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Raphael Morard

Dr. Raphael Morard

Group:	Micropaleontology - Paleoceanography
Phone:	+49 421 218-65979
Email:	[Bitte aktivieren Sie Javascript]
Room:	MARUM II, room 1040
Address:	Universität Bremen MARUM – Zentrum für Marine Umweltwissenschaften Leobener Str. 8 D-28359 Bremen
External Profiles:	OrcID \| Google Scholar \| Research Gate

Research interests

I am a micropaleontologist and molecular ecologist focused on the biodiversity, ecology, and evolution of foraminifera, particularly in their use as indicators of past marine environments. In my research, I use foraminifera as a model group to investigate broader patterns of marine evolution. By combining molecular techniques, such as molecular taxonomy and metagenomics, with quantitative micropaleontology and morphometry, I aim to understand how marine species evolve, adapt, interact with other species, and spread to new areas. My research aims to gain insights into how marine life responds to global environmental changes.

Cryptic diversity in plankton

My earliest research interest is to investigate how new species emerge in the open ocean. This environment is largely unstructured and nutrient-poor, yet it supports surprisingly diverse ecosystems—a phenomenon known as the "plankton paradox". Part of this diversity is "cryptic diversity"—or hidden diversity—where nearly identical species are genetically distinct. This indicates that much of the diversification process is not visible through traditional morphology-based methods. I am using planktonic foraminifera as model organisms and employing a combination of barcoding and metabarcoding methodologies in my research to better understand how diversity is structured and maintained at this cryptic level.

Living planktonic foraminifera — Living assemblage of planktonic foraminifera (©Michael Siccha)

Abbildung zeigt — Invaded environment in the mediterranean sea at Capo Passero, Sicily (©Débora Raposo)

Biological Invasion
I am also interested in exploring the process of tropicalization in the Mediterranean Sea. This phenomenon involves the transformation of ecosystems as tropical and subtropical species expand toward the poles, while temperate species retract, largely due to climate change. In the Mediterranean, this shift began with the opening of the Suez Canal in 1869, which removed a natural biogeographic barrier and allowed a largely one-way influx of Indo-Pacific marine species. To date, over 600 invasive species have been documented, and the process is expected to be amplified by climate change. In my research, I use benthic foraminifera as a model to examine the factors that lead to successful invasions, with a focus on population structure, symbiosis, and tolerance to varying temperature extremes.

Ancient DNA
I am captivated by the emerging field of ancient DNA (aDNA) in marine environments. While aDNA has been widely used to reconstruct past terrestrial ecosystems, its application in marine contexts is still in the early stages. Recent advancements in DNA extraction and sequencing technologies now make it possible to recover the DNA of ancient marine communities from sediments dating back several hundred thousand years. This approach offers vast potential for paleoceanography and paleoecology, as it provides insights beyond those limited to organisms with fossilized remains—a small fraction of past biodiversity. With ancient DNA, we can reconstruct entire ecosystems and understand how they transformed under various conditions, including some that no longer exist today.

Expeditions

MsM111 - F/S Merian	North Atlantic and Baffin Bay (Reikjavik, Iceland - St John’s, Canada, September 2022)
M140 - F/S Meteor	Subtropical Eastern Atlantic (Mindele, Cabo Verde - Las Palmas,Gran Canaria, August 2017)
M124 - F/S Meteor	Subtropical South Atlantic (Cap Town, South Africa - Rio de Janeiro, Brazil, March 2016)
MsM39 - F/S Merian	North West Atlantic (St John's - St John's, Canada, June 2014)
DipFIP - F/S SONNE	Western Equatorial Pacific (Wellington, New Zealand - Kaoshiung, Taiwan, March 2013)
Tara Ocean expedition - TARA schooner	North East Pacific (Hawaï - San Diego, USA, Sept. - Oct. 2011).
FORCLIM-7 - R/V Sarmiento de Gamboa	Gulf of Biscay (Vigo – Vigo, Spain, April 2009).
GYRAFOR-A - R/V ALIS	Western equatorial Pacific (Nouméa – Nouméa, New Caledonia, Jun.-Jul. 2008).

Teaching at the University of Bremen

Since 2024	Ecology for the Natural Sciences for Sustainability bachelor
Since 2024	Quantitative Methods in Paleontology for the Marine Geosciences bachelor
Since 2023	Earth and Life History for the Marine Geosciences bachelor
Since 2022	Marine Micropaleontology for the Marine Geosciences bachelor
Since 2022	Eukaryotic Microbiology for the MarMic Master
Since 2020	Mapping course in Vorarlberg (Austria) for the Geowissenschaften bachelor
2018-2023	Paleoecology for the Geowissenschaften bachelor

CV

Research positions

Since 2021	Senior scientist at MARUM, University of Bremen.
2013-2021	Post-Doctoral researcher at MARUM, University of Bremen.
2011-2012	Post-Doctoral researcher at Biological station of Roscoff.
2007-2010	PhD Student at University Claude Bernard Lyon I.

Education

2007 - 2010	PhD degree in Micropaleontology. University Claude Bernard Lyon I, France
2005 - 2007	Master degree in Paleontology, Sedimentology and Paleoenvironment. University Claude Bernard Lyon I
2002 - 2005	Bachelor degree in Earth sciences. University Claude Bernard Lyon I

Publication List

2024

Girard, E.B., Didaskalou, E.A., Pratama, A.M.A., Rattner, C., Morard, R., Renema, W., 2024. Quantitative assessment of reef foraminifera community from metabarcoding data. Mol Ecol Resour. https://doi.org/10.1111/1755-0998.14000

Manda, S., Herut, B., Rilov, G., Kucera, M., Morard, R., Abramovich, S., Ashckenazi-Polivoda, S., 2024. A dynamic subtropical coastal hotspot of benthic foraminifera in the Southeastern Mediterranean indicates early-stage tropicalization. Science of The Total Environment 945, 173917. https://doi.org/10.1016/j.scitotenv.2024.173917

Meilland, J., Siccha, M., Morard, R., Kucera, M., 2024. Continuous reproduction in laboratory culture of planktonic foraminifera. Eukaryotic Microbiology 1–16. https://doi.org/10.1111/jeu.13022

Morard, R., Darling, K.F., Weiner, A.K.M., Hassenrück, C., Vanni, C., Cordier, T., Henry, N., Greco, M., Vollmar, N.M., Milivojević, T., Rahman, S.N., Siccha, M., Meilland, J., Jonkers, L., Quillévéré, F., Escarguel, G., Douady, C.J., de Garidel-Thoron, T., de Vargas, C., Kucera, M., 2024. The global genetic diversity of planktonic foraminifera reveals the structure of cryptic speciation in plankton. Biological Reviews. https://doi.org/10.1111/brv.13065

2023

Greco, M., Morard, R., Darling, K., Kucera, M., 2023. Macroevolutionary patterns in intragenomic rDNA variability among planktonic foraminifera. PeerJ 11, e15255. https://doi.org/10.7717/peerj.15255

Meilland, J., Ezat, M.M., Westgård, A., Manno, C., Morard, R., Siccha, M., Kucera, M., 2023. Rare but persistent asexual reproduction explains the success of planktonic foraminifera in polar oceans. J Plankton Res 45, 15–32. https://doi.org/10.1093/plankt/fbac069

Silva Raposo, D., A. Zufall, R., Caruso, A., Titelboim, D., Abramovich, S., Hassenrück, C., Kucera, M., Morard, R., 2023. Invasion success of a Lessepsian symbiont-bearing foraminifera linked to high dispersal ability, preadaptation and suppression of sexual reproduction. Sci Rep 13, 12578. https://doi.org/10.1038/s41598-023-39652-y

Siccha, M., Morard, R., Meilland, J., Iwasaki, S., Kucera, M., Kimoto, K., 2023. Collection of X-ray micro computed tomography images of shells of planktic foraminifera with curated taxonomy. Sci Data 10, 679. https://doi.org/10.1038/s41597-023-02498-0

Pinko, D., Abramovich, S., Rahav, E., Belkin, N., Rubin-blum, M., Kucera, M., Morard, R., Holzmann, M., Abdu, U., 2023. Shared ancestry of algal symbiosis and chloroplast sequestration in foraminifera 3401. https://doi.org/10.1126/sciadv.adi3401

2022

Cordier, T., Angeles, I.B., Henry, N., Lejzerowicz, F., Berney, C., Morard, R., Brandt, A., Cambon-Bonavita, M.-A., Guidi, L., Lombard, F., Arbizu, P.M., Massana, R., Orejas, C., Poulain, J., Smith, C.R., Wincker, P., Arnaud-Haond, S., Gooday, A.J., de Vargas, C., Pawlowski, J., 2022. Patterns of eukaryotic diversity from the surface to the deep-ocean sediment. Sci Adv 8, 1–13. https://doi.org/10.1126/sciadv.abj9309

Holzmann, M., Hohenegger, J., Apothéloz-Perret-Gentil, L., Morard, R., Abramovich, S., Titelboim, D., Pawlowski, J., 2022. Operculina and Neoassilina: A Revision of Recent Nummulitid Genera Based on Molecular and Morphological Data Reveals a New Genus. Journal of Earth Science 33, 1411–1424. https://doi.org/10.1007/s12583-021-1595-8

Meilland, J., Cornuault, P., Morard, R., Brummer, G.A., Kucera, M., 2022. Identification guide to extant planktonic foraminifera. Part 1: Family Candeinidae and genera Berggrenia, Bolivina, Dentigloborotalia, and Neogallitellia. ICES Identification leaflets for Plankton No. 196, 22. https://doi.org/https://doi.org/10.17895/ices.pub.7643

Morard, R., Hassenrück, C., Greco, M., Fernandez-Guerra, A., Rigaud, S., Douady, C.J., Kucera, M., 2022. Renewal of planktonic foraminifera diversity after the Cretaceous Paleogene mass extinction by benthic colonizers. Nat Commun 13, 1–9. https://doi.org/10.1038/s41467-022-34794-5

2021

Greco, M., Morard, R., Kucera, M., 2021. Single-cell metabarcoding reveals biotic interactions of the Arctic calcifier Neogloboquadrina pachyderma with the eukaryotic pelagic community. J Plankton Res 43, 113–125. https://doi.org/10.1093/plankt/fbab015

Jonkers, L., Gopalakrishnan, A., Weßel, L., Chiessi, C.M., Groeneveld, J., Monien, P., Lessa, D., Morard, R., 2021. Morphotype and Crust Effects on the Geochemistry of Globorotalia inflata . Paleoceanogr Paleoclimatol 36, 1–16. https://doi.org/10.1029/2021pa004224

Lejzerowicz, F., Gooday, A.J., Barrenechea Angeles, I., Cordier, T., Morard, R., Apothéloz-Perret-Gentil, L., Lins, L., Menot, L., Brandt, A., Levin, L.A., Martinez Arbizu, P., Smith, C.R., Pawlowski, J., 2021. Eukaryotic Biodiversity and Spatial Patterns in the Clarion-Clipperton Zone and Other Abyssal Regions: Insights From Sediment DNA and RNA Metabarcoding. Front Mar Sci 8, 1–23. https://doi.org/10.3389/fmars.2021.671033

Titelboim, D., Thangarjan, S., Raposo, D., Morard, R., Kucera, M., Ashckenazi‐Polivoda, S., Almogi‐Labin, A., Herut, B., Manda, S., Abramovich, S., Gold, D.A., Abdu, U., 2021. The transcriptomic signature of cold and heat stress in benthic foraminifera—Implications for range expansions of marine calcifiers. Funct Ecol 35, 2679–2690. https://doi.org/10.1111/1365-2435.13929

Milivojević, T., Rahman, S.N., Raposo, D., Siccha, M., Kucera, M., Morard, R., 2021. High variability in SSU rDNA gene copy number among planktonic foraminifera revealed by single-cell qPCR. ISME Communications 1, 63. https://doi.org/10.1038/s43705-021-00067-3

2020

Barrenechea Angeles, I., Lejzerowicz, F., Cordier, T., Scheplitz, J., Kucera, M., Ariztegui, D., Pawlowski, J., Morard, R., 2020. Planktonic foraminifera eDNA signature deposited on the seafloor remains preserved after burial in marine sediments. Sci. Rep. 10, 1–12. doi:10.1038/s41598-020-77179-8

Lessa, D., Morard, R., Jonkers, L., M. Venancio, I., Reuter, R., Baumeister, A., Luiza Albuquerque, A., Kucera, M., 2020. Distribution of planktonic foraminifera in the subtropical South Atlantic: Depth hierarchy of controlling factors. Biogeosciences 17. doi:10.5194/bg-17-4313-2020

Li, Q., Lei, Y., Morard, R., Li, T., Wang, B., 2020. Diversity hotspot and unique community structure of foraminifera in the world’s deepest marine blue hole – Sansha Yongle Blue Hole. Sci. Rep. 1–11. doi:10.1038/s41598-020-67221-0

Orsi, W.D., Morard, R., Vuillemin, A., Eitel, M., Wörheide, G., Milucka, J., Kucera, M., 2020. Anaerobic metabolism of Foraminifera thriving below the seafloor. ISME J. doi:10.1038/s41396-020-0708-1

Prazeres, M., Morard, R., Roberts, T.E., Doo, S.S., Jompa, J., Schmidt, C., Stuhr, M., Renema, W., Kucera, M., 2020. High dispersal capacity and biogeographic breaks shape the genetic diversity of a globally distributed reef‐dwelling calcifier. Ecol. Evol. 10, 5976–5989. doi:10.1002/ece3.6335

2019

Meilland, J., Siccha, M., Weinkauf, M.F.G., Jonkers, L., Morard, R., Baranowski, U., Baumeister, A., Bertlich, J., Brummer, G.-J., Debray, P., Fritz-Endres, T., Groeneveld, J., Magerl, L., Munz, P., Rillo, M.C., Schmidt, C., Takagi, H., Theara, G., Kucera, M., 2019. Highly replicated sampling reveals no diurnal vertical migration but stable species-specific vertical habitats in planktonic foraminifera. J. Plankton Res. 41. doi:10.1093/plankt/fbz002

Morard, R., Vollmar, N.M., Greco, M., Kucera, M., 2019. Unassigned diversity of planktonic foraminifera from environmental sequencing revealed as known but neglected species. doi:10.1371/journal.pone.0213936

Morard, R., Füllberg, A., Brummer, G.J.A., Greco, M., Jonkers, L., Wizemann, A., Weiner, A.K.M., Darling, K., Siccha, M., Ledevin, R., Kitazato, H., De Garidel-Thoron, T., De Vargas, C., Kucera, M., 2019. Genetic and morphological divergence in the warm-water planktonic foraminifera genus Globigerinoides. PLoS One 14, 1–30. doi:10.1371/journal.pone.0225246

2018

André, A., Quillévéré, F., Schiebel, R., Morard, R., Howa, H., Meilland, J., Douady, C.J., 2018. Disconnection between genetic and morphological diversity in the planktonic foraminifer Neogloboquadrina pachyderma from the Indian sector of the Southern Ocean. Mar. Micropaleontol. 144, 14–24. doi:10.1016/j.marmicro.2018.10.001

Morard, R., Mahé, F., Romac, S., Poulain, J., Kucera, M., Vargas, C. De, 2018. Surface ocean metabarcoding confirms limited diversity in planktonic foraminifera but reveals unknown hyper-abundant lineages. Sci. Rep. 8:2539, 1–10. doi:10.1038/s41598-018-20833-z

Schiebel, R., Smart, S.M., Jentzen, A., Jonkers, L., Morard, R., Meilland, J., Michel, E., Coxall, H.K., Hull, P.M., de Garidel-Thoron, T., Aze, T., Quillévéré, F., Ren, H., Sigman, D.M., Vonhof, H.B., Martínez-García, A., Kučera, M., Bijma, J., Spero, H.J., Haug, G.H., 2018. Advances in planktonic foraminifer research: New perspectives for paleoceanography. Rev. Micropaléontologie 1–26. doi:10.1016/J.REVMIC.2018.10.001

Schmidt, C., Morard, R., Romero, O., Kucera, M., 2018. Diverse Internal Symbiont Community in the Endosymbiotic Foraminifera Pararotalia calcariformata: Implications for Symbiont Shuffling Under Thermal Stress. Front. Microbiol. 9, 1–11. doi:10.3389/fmicb.2018.02018

2017

Morard, R., Lejzerowicz, F., Darling, K.F., Lecroq-bennet, B., Pedersen, M.W., Orlando, L., Pawlowski, J., Mulitza, S., Vargas, C. De, Kucera, M., 2017. Planktonic foraminifera-derived environmental DNA extracted from abyssal sediments preserves patterns of plankton macroecology. Biogeosciences 14, 2741–2754. doi:10.5194/bg-14-2741-2017

Kucera, M., Silye, L., Weiner, A.K.M., Darling, K.F., Lübben, B., Holzmann, M., Pawlowski, J., Schönfeld, J., Morard, R., 2017. Caught in the act: anatomy of an ongoing benthic–planktonic transition in a marine protist. J. Plankton Res. 39, 436–449. doi:10.1093/plankt/fbx018

2016

Morard, R., Escarguel, G., Weiner, A.K.M., André, A., Douady, C.J., Wade, C.M., Darling, K.F., Ujiié, Y., Seears, H.A., Quillévéré, F., de Garidel-Thoron, T., de Vargas, C., Kucera, M., 2016. Nomenclature for the Nameless: A Proposal for an Integrative Molecular Taxonomy of Cryptic Diversity Exemplified by Planktonic Foraminifera. Syst. Biol. 65, 925–940. doi:10.1093/sysbio/syw031

Morard, R., Reinelt, M., Chiessi, C.M., Groeneveld, J., Kucera, M., 2016. Tracing shifts of oceanic fronts using the cryptic diversity of the planktonic foraminifera Globorotalia inflata. Paleoceanography 31, 1193–1205. doi:10.1002/2016PA002977

Schmidt, C., Morard, R., Prazeres, M., Barak, H., Kucera, M., 2016. Retention of high thermal tolerance in the invasive foraminifera Amphistegina lobifera from the Eastern Mediterranean and the Gulf of Aqaba. Mar. Biol. 163, 228. doi:10.1007/s00227-016-2998-4

Weiner, A.K.M., Morard, R., Weinkauf, M.F., Darling, K.F., André, A., Quillévéré, F., Ujiie, Y., Douady, C.J., de Vargas, C., Kucera, M., 2016. Methodology for single-cell genetic analysis of planktonic foraminifera for studies of protist diversity and evolution. Fontiers Mar. Sci. 3, 1–15. doi:10.3389/fmars.2016.00255

2015

Schmidt, C., Morard, R., Almogi-Labin, A., Weinmann, A.E., Titelboim, D., Abramovich, S., Kucera, M., 2015. Recent Invasion of the Symbiont-Bearing Foraminifera Pararotalia into the Eastern Mediterranean Facilitated by the Ongoing Warming Trend. PLoS One 10, e0132917. doi:10.1371/journal.pone.0132917

Spezzaferri, S., Kucera, M., Pearson, P.N., Wade, B.S., Rappo, S., Poole, C.R., Morard, R., Stalder, C., 2015. Fossil and genetic evidence for the polyphyletic nature of the planktonic foraminifera “Globigerinoides”, and description of the new genus Trilobatus. PLoS One 10, e0128108. doi:10.1371/journal.pone.0128108

De Vargas, C., Audic, S., Henry, N., Decelle, J., Mahe, F., Logares, R., Lara, E., Berney, C., Le Bescot, N., Probert, I., Carmichael, M., Poulain, J., Romac, S., Colin, S., Aury, J.-M., Bittner, L., Chaffron, S., Dunthorn, M., Engelen, S., Flegontova, O., Guidi, L., Horak, A., Jaillon, O., Lima-Mendez, G., Luke, J., Malviya, S., Morard, R., Mulot, M., Scalco, E., Siano, R., Vincent, F., Zingone, A., Dimier, C., Picheral, M., Searson, S., Kandels-Lewis, S., Acinas, S.G., Bork, P., Bowler, C., Gorsky, G., Grimsley, N., Hingamp, P., Iudicone, D., Not, F., Ogata, H., Pesant, S., Raes, J., Sieracki, M.E., Speich, S., Stemmann, L., Sunagawa, S., Weissenbach, J., Wincker, P., Karsenti, E., Boss, E., Follows, M., Karp-Boss, L., Krzic, U., Reynaud, E.G., Sardet, C., Sullivan, M.B., Velayoudon, D., 2015. Eukaryotic plankton diversity in the sunlit ocean. Science. 348, 1–11. doi:10.1126/science.1261605

Morard, R., Darling, K.F., Mahé, F., Audic, S., Ujiié, Y., Weiner, A.K.M., André, A., Seears, H. a., Wade, C.M., Quillévéré, F., Douady, C.J., Escarguel, G., de Garidel-Thoron, T., Siccha, M., Kucera, M., de Vargas, C., 2015. PFR2: a curated database of planktonic Foraminifera18S ribosomal DNA as a resource for studies of plankton ecology, biogeography, and evolution. Mol. Ecol. Resour. 49, 1–14. doi:10.1111/1755-0998.12410

2014

André, A., Quillévéré, F., Morard, R., Ujiié, Y., Escarguel, G., de Vargas, C., de Garidel-Thoron, T., Douady, C.J., 2014. SSU rDNA Divergence in Planktonic Foraminifera: Molecular Taxonomy and Biogeographic Implications. PLoS One 9, e104641. doi:10.1371/journal.pone.0104641

2013

Morard, R., Quillévéré, F., Escarguel, G., Garidel-thoron, T. De, 2013. Ecological modeling of the temperature dependence of cryptic species of planktonic Foraminifera in the Southern Hemisphere. Palaeogeography, Palaeoclimatology, Palaeocology. 391 Part A, 13–33.

Quilllévéré, F., Morard, R., Escarguel, G., Douady, C.J., Ujiié, Y., de Garidel-Thoron, T., de Vargas, C. 2013. Global scale same-specimen morpho-genetic analysis of Truncorotalia truncatulinoides: A perspective on the morphological species concept in planktonic foraminifera. Palaeogeography, Palaeoclimatology, Palaeocology, 391 Part A, 2–12.

André, A., Weiner, A., Quillévéré, F., Aurahs, R., Morard, R., Douady, C.J., Garidel-Thoron, T. de, Escarguel, G., Vargas, C. de, Kucera, M., 2013. The cryptic and the apparent reversed : lack of genetic differentiation within the morphologically diverse plexus of the planktonic foraminifer Globigerinoides sacculifer. Paleobiology 39, 21–39.

Guillou, L., Bachar, D., Audic, S., Bass, D., Berney, C., Bittner, L., Boutte, C., Burgaud, G., de Vargas, C., Decelle, J., Del Campo, J., Dolan, J.R., Dunthorn, M., Edvardsen, B., Holzmann, M., Kooistra, W.H.C.F., Lara, E., Le Bescot, N., Logares, R., Mahé, F., Massana, R., Montresor, M., Morard, R., Not, F., Pawlowski, J., Probert, I., Sauvadet, A.-L., Siano, R., Stoeck, T., Vaulot, D., Zimmermann, P., Christen, R., 2013. The Protist Ribosomal Reference database (PR2): a catalog of unicellular eukaryote small sub-unit rRNA sequences with curated taxonomy. Nucleic Acids Res. 41, D597–604.

2011

Morard, R., Quillévéré, F., Douady, C.J., de Vargas, C., de Garidel-Thoron, T., Escarguel, G., 2011. Worldwide genotyping in the planktonic foraminifer Globoconella inflata: Implications for life history and paleoceanography. PLoS One 6, e26665. doi:10.1371/journal.pone.0026665

2009

Morard, R., Quillévéré, F., Escarguel, G., Ujiie, Y., de Garidel-Thoron, T., Norris, R.D., de Vargas, C., 2009. Morphological recognition of cryptic species in the planktonic foraminifer Orbulina universa.Mar. Micropaleontol. 71, 148–165. doi:10.1016/j.marmicro.2009.03.001

Contribution to Academic conferences and seminars

Morard, R. Constraining the biodiversity of planktonic foraminifera using Next-Generation Sequencing. Eukaryotic-Omics, Geneva , Switzerland, 25-26th April 2014.
Morard, R. Constraining the biodiversity of marine microplankton using Next-Generation Sequencing. Biology Seminar, Kaiserslautern, Germany, 3rd of February 2014.
Weiner, A., Morard, R., Kucera, M., (2014) Extent of cryptic diversity in planktonic foraminifera morphospecies. http://www.sgmeet.com/osm2014/viewabstract.asp?AbstractID=12992. Ocean Sciences Meeting 2014, Honolulu, USA).
Weiner, A., Morard, R., Kucera, M. Genetic diversity and phylogeography of planktonic Foraminifera. In: Qian PY, Kolter R (eds) Gordon Research Conference on Marine Molecular Ecology 2013, Hongkong, 2013.
Morard, R., Mahé, F., Audic, S., Romac, S., Ujiié, Y., Garet-Delmas, M. J., Weiner, A., Kucera, M., de Vargas, C. The potential of Next Generation Sequencing to constrain the global biodiversity of planktonic foraminifera. The Micropalaeontological Society’s Spring Meeting, Prague, Czech Republic, June 19th-22, 2013
Morard R., Le Bescot, N. Mahé, F., Audic S., Romac S., Not F., Ujiié Y., de Vargas C. Reaching the boundaries of global planktonic foraminiferal biodiversity using Next Generation Sequencing. SCOR/IGBP Working group 138, Modern Planktic Foraminifera and Ocean changes, Amsterdam (Netherlands), 1st September 2011.
Morard R., Quillévéré F., Douady C. J., Escarguel G., de Garidel-Thoron T and de Vargas C. The allopatric distribution of Globoconella inflata cryptic species and its potential used for monitoring past movements of the Antarctic subpolar front. FORAMS 2010, Bonn, Germany, September 5th-10th, 2010.
Morard, R. Morphometric analyses of planktonic foraminiferal cryptic species: new perspectives for paleoceanographic reconstructions. Seminar, Tuebingen. 20th of June 2010.
Quillévéré F., Morard R., de Vargas C. Douady C. J., Ujiié Y., de Garidel-Thoron T and Escarguel G. Truncorotalia truncatulinoides and the value of direct morpho-genetic comparisons in planktonic foraminifera. FORAMS 2010, Bonn, Germany, September 5th-10th, 2010.
Morard, R., Quillévéré, F., Escarguel, G., Ujiié, Y., De Garidel Thoron T., Lécuyer C. and de Vargas C. Morphometric and stable isotope (δ13C, δ18O) analyses of planktonic foraminiferal cryptic species: new perspectives for paleoceanographic reconstructions. Bioindicators of past and present environments, The Micropalaeontological Society’s Spring Meeting, Tübingen, Germany, 15th -17th May 2008.
Morard, R. Eco- morpho- genotypes in planktonic foraminifera : new perspectives for paleoceanographic reconstructions. Sarco – Deal, Villefranche sur Mer, 13-15th November 2007.

Cruise reports

Kučera, M., Faust, J. C., Meilland, J., Dmello, G., Janßen, A., Schulz, H., Morard, R., Streuff, K., Vernal, A. De, Düßmann, R., Bergenthal, M., Linowski, E., Fröhlich, S., Ahrlich, F., Schillai, S., Schmidt, W., Haider, D., Detlef, H., Diekamp, V., & Siccha, M. (2023). Deep drilling in the Baffin Bay for marine sediment records of Greenland Ice Sheet collapse during past warm intervals. https://doi.org/10.48433/cr_msm111.

Kucera, M., Siccha, M., Morard, R., Jonkers, L., Schmidt, C., Munz, P., Groeneveld, J., Fischer, G., Ruhland, G., Klann, M., Iversen, M., Konrad, C., Stuut, J.-B., Brummer, G.-J., Koster, B., Witte, Y., Visser, J.-D. de, Weinkauf, M., Takagi, H., … Raeke, A. (2019). Scales of Population Dynamics, Ecology and Diversity of Planktonic Foraminifera and their Relationship to Particle Flux in the Eastern Tropical Atlantic: Cruise No. M140, 11.8.2017-5.9.2017, Mindelo (Cabo Verde)-Las Palmas (Spain) - FORAMFLUX. https://doi.org/doi: 10.2312/cr_m140.

Kucera, M., Morard, R., Siccha, M., Weiner, A., & Weinkauf, M. (2013). Cruise report of RV Sonne Cruise SO226-3 DipFIP ‒ the extent and structure of cryptic diversity in morphospecies of planktonic Foraminifera of the Indopacific Warm Pool: Wellington‒Kaohsiung, 04.03.2013‒28.03.2013 (Issue 293).